In HU this root, originally meaning "gullet; intestines", shifted the meaning and yielded *tis- (Hurr. tiža, tiž-na, Ur. tiš-nu) "heart" (after the basic word for "heart", *jĕrḳwi, changed its meaning to "middle, inside"). See Diakonoff-Starostin 1986, 53 (with erroneous attraction of Rut. lɨs).
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